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**EXPERIMENTAL** CallableLoci

Determine callable status of loci

Category Coverage Analysis


Overview

Collect statistics on callable, uncallable, poorly mapped, and other parts of the genome

A very common question about a NGS set of reads is what areas of the genome are considered callable. This tool considers the coverage at each locus and emits either a per base state or a summary interval BED file that partitions the genomic intervals into the following callable states:

REF_N
The reference base was an N, which is not considered callable the GATK
PASS
The base satisfied the min. depth for calling but had less than maxDepth to avoid having EXCESSIVE_COVERAGE
NO_COVERAGE
Absolutely no reads were seen at this locus, regardless of the filtering parameters
LOW_COVERAGE
There were fewer than min. depth bases at the locus, after applying filters
EXCESSIVE_COVERAGE
More than --max-depth read at the locus, indicating some sort of mapping problem
POOR_MAPPING_QUALITY
More than --max-fraction-of-reads-with-low-mapq at the locus, indicating a poor mapping quality of the reads

Input

A BAM file containing exactly one sample.

Output

A file with the callable status covering each base and a table of callable status x count of all examined bases

Usage example

  gatk CallableLoci \
      -I myreads.bam \
      -R myreference.fasta \
      -O callable_status.bed \    
      --summary table.txt 
 

would produce a BED file that looks like:

     20 10000000 10000864 PASS
     20 10000865 10000985 POOR_MAPPING_QUALITY
     20 10000986 10001138 PASS
     20 10001139 10001254 POOR_MAPPING_QUALITY
     20 10001255 10012255 PASS
     20 10012256 10012259 POOR_MAPPING_QUALITY
     20 10012260 10012263 PASS
     20 10012264 10012328 POOR_MAPPING_QUALITY
     20 10012329 10012550 PASS
     20 10012551 10012551 LOW_COVERAGE
     20 10012552 10012554 PASS
     20 10012555 10012557 LOW_COVERAGE
     20 10012558 10012558 PASS
 
as well as a summary table that looks like:

                        state nBases
                        REF_N 0
                         PASS 996046
                  NO_COVERAGE 121
                 LOW_COVERAGE 928
           EXCESSIVE_COVERAGE 0
         POOR_MAPPING_QUALITY 2906
 
@author Mark DePristo / Jonn Smith @since May 7, 2010 / Nov 1, 2024

Additional Information

Read filters

These Read Filters are automatically applied to the data by the Engine before processing by CallableLoci.

CallableLoci specific arguments

This table summarizes the command-line arguments that are specific to this tool. For more details on each argument, see the list further down below the table or click on an argument name to jump directly to that entry in the list.

Argument name(s) Default value Summary
Required Arguments
--input
 -I
BAM/SAM/CRAM file containing reads
--output
 -O
Output file (BED or per-base format)
--reference
 -R
Reference sequence file
--summary
Name of file for output summary
Optional Tool Arguments
--arguments_file
read one or more arguments files and add them to the command line
--cloud-index-prefetch-buffer
 -CIPB
-1 Size of the cloud-only prefetch buffer (in MB; 0 to disable). Defaults to cloudPrefetchBuffer if unset.
--cloud-prefetch-buffer
 -CPB
40 Size of the cloud-only prefetch buffer (in MB; 0 to disable).
--disable-bam-index-caching
 -DBIC
false If true, don't cache bam indexes, this will reduce memory requirements but may harm performance if many intervals are specified. Caching is automatically disabled if there are no intervals specified.
--disable-sequence-dictionary-validation
false If specified, do not check the sequence dictionaries from our inputs for compatibility. Use at your own risk!
--gcs-max-retries
 -gcs-retries
20 If the GCS bucket channel errors out, how many times it will attempt to re-initiate the connection
--gcs-project-for-requester-pays
Project to bill when accessing "requester pays" buckets. If unset, these buckets cannot be accessed. User must have storage.buckets.get permission on the bucket being accessed.
--help
 -h
false display the help message
--interval-merging-rule
 -imr
ALL Interval merging rule for abutting intervals
--intervals
 -L
One or more genomic intervals over which to operate
--max-depth
Maximum read depth before a locus is considered poorly mapped
--max-depth-per-sample
0 Maximum number of reads to retain per sample per locus. Reads above this threshold will be downsampled. Set to 0 to disable.
--max-fraction-of-reads-with-low-mapq
 -frlmq
0.1 If the fraction of reads at a base with low mapping quality exceeds this value, the site may be poorly mapped
--max-low-mapq
 -mlmq
1 Maximum value for MAPQ to be considered a problematic mapped read
--min-base-quality
 -mbq
20 Minimum quality of bases to count towards depth
--min-mapping-quality
 -mmq
10 Minimum mapping quality of reads to count towards depth
--sites-only-vcf-output
false If true, don't emit genotype fields when writing vcf file output.
--version
false display the version number for this tool
Optional Common Arguments
--add-output-sam-program-record
true If true, adds a PG tag to created SAM/BAM/CRAM files.
--add-output-vcf-command-line
true If true, adds a command line header line to created VCF files.
--create-output-bam-index
 -OBI
true If true, create a BAM/CRAM index when writing a coordinate-sorted BAM/CRAM file.
--create-output-bam-md5
 -OBM
false If true, create a MD5 digest for any BAM/SAM/CRAM file created
--create-output-variant-index
 -OVI
true If true, create a VCF index when writing a coordinate-sorted VCF file.
--create-output-variant-md5
 -OVM
false If true, create a a MD5 digest any VCF file created.
--disable-read-filter
 -DF
Read filters to be disabled before analysis
--disable-tool-default-read-filters
false Disable all tool default read filters (WARNING: many tools will not function correctly without their default read filters on)
--exclude-intervals
 -XL
One or more genomic intervals to exclude from processing
--gatk-config-file
A configuration file to use with the GATK.
--interval-exclusion-padding
 -ixp
0 Amount of padding (in bp) to add to each interval you are excluding.
--interval-padding
 -ip
0 Amount of padding (in bp) to add to each interval you are including.
--interval-set-rule
 -isr
UNION Set merging approach to use for combining interval inputs
--inverted-read-filter
 -XRF
Inverted (with flipped acceptance/failure conditions) read filters applied before analysis (after regular read filters).
--lenient
 -LE
false Lenient processing of VCF files
--max-variants-per-shard
0 If non-zero, partitions VCF output into shards, each containing up to the given number of records.
--QUIET
false Whether to suppress job-summary info on System.err.
--read-filter
 -RF
Read filters to be applied before analysis
--read-index
Indices to use for the read inputs. If specified, an index must be provided for every read input and in the same order as the read inputs. If this argument is not specified, the path to the index for each input will be inferred automatically.
--read-validation-stringency
 -VS
SILENT Validation stringency for all SAM/BAM/CRAM/SRA files read by this program. The default stringency value SILENT can improve performance when processing a BAM file in which variable-length data (read, qualities, tags) do not otherwise need to be decoded.
--seconds-between-progress-updates
10.0 Output traversal statistics every time this many seconds elapse
--sequence-dictionary
Use the given sequence dictionary as the master/canonical sequence dictionary. Must be a .dict file.
--tmp-dir
Temp directory to use.
--use-jdk-deflater
 -jdk-deflater
false Whether to use the JdkDeflater (as opposed to IntelDeflater)
--use-jdk-inflater
 -jdk-inflater
false Whether to use the JdkInflater (as opposed to IntelInflater)
--verbosity
INFO Control verbosity of logging.
Advanced Arguments
--format
BED Output format
--min-depth
4 Minimum QC+ read depth before a locus is considered callable
--min-depth-for-low-mapq
 -mdflmq
10 Minimum read depth before a locus is considered a potential candidate for poorly mapped
--showHidden
false display hidden arguments

Argument details

Arguments in this list are specific to this tool. Keep in mind that other arguments are available that are shared with other tools (e.g. command-line GATK arguments); see Inherited arguments above.


--add-output-sam-program-record / -add-output-sam-program-record

If true, adds a PG tag to created SAM/BAM/CRAM files.

boolean  true


--add-output-vcf-command-line / -add-output-vcf-command-line

If true, adds a command line header line to created VCF files.

boolean  true


--arguments_file

read one or more arguments files and add them to the command line

List[File]  []


--cloud-index-prefetch-buffer / -CIPB

Size of the cloud-only prefetch buffer (in MB; 0 to disable). Defaults to cloudPrefetchBuffer if unset.

int  -1  [ [ -∞  ∞ ] ]


--cloud-prefetch-buffer / -CPB

Size of the cloud-only prefetch buffer (in MB; 0 to disable).

int  40  [ [ -∞  ∞ ] ]


--create-output-bam-index / -OBI

If true, create a BAM/CRAM index when writing a coordinate-sorted BAM/CRAM file.

boolean  true


--create-output-bam-md5 / -OBM

If true, create a MD5 digest for any BAM/SAM/CRAM file created

boolean  false


--create-output-variant-index / -OVI

If true, create a VCF index when writing a coordinate-sorted VCF file.

boolean  true


--create-output-variant-md5 / -OVM

If true, create a a MD5 digest any VCF file created.

boolean  false


--disable-bam-index-caching / -DBIC

If true, don't cache bam indexes, this will reduce memory requirements but may harm performance if many intervals are specified. Caching is automatically disabled if there are no intervals specified.

boolean  false


--disable-read-filter / -DF

Read filters to be disabled before analysis

List[String]  []


--disable-sequence-dictionary-validation / -disable-sequence-dictionary-validation

If specified, do not check the sequence dictionaries from our inputs for compatibility. Use at your own risk!

boolean  false


--disable-tool-default-read-filters / -disable-tool-default-read-filters

Disable all tool default read filters (WARNING: many tools will not function correctly without their default read filters on)

boolean  false


--exclude-intervals / -XL

One or more genomic intervals to exclude from processing
Use this argument to exclude certain parts of the genome from the analysis (like -L, but the opposite). This argument can be specified multiple times. You can use samtools-style intervals either explicitly on the command line (e.g. -XL 1 or -XL 1:100-200) or by loading in a file containing a list of intervals (e.g. -XL myFile.intervals). strings gathered from the command line -XL argument to be parsed into intervals to exclude

List[String]  []


--format / -format

Output format
The output of this tool will be written in this format. The recommended option is BED.

The --format argument is an enumerated type (OutputFormat), which can have one of the following values:

BED
STATE_PER_BASE

OutputFormat  BED


--gatk-config-file

A configuration file to use with the GATK.

String  null


--gcs-max-retries / -gcs-retries

If the GCS bucket channel errors out, how many times it will attempt to re-initiate the connection

int  20  [ [ -∞  ∞ ] ]


--gcs-project-for-requester-pays

Project to bill when accessing "requester pays" buckets. If unset, these buckets cannot be accessed. User must have storage.buckets.get permission on the bucket being accessed.

String  ""


--help / -h

display the help message

boolean  false


--input / -I

BAM/SAM/CRAM file containing reads

R List[GATKPath]  []


--interval-exclusion-padding / -ixp

Amount of padding (in bp) to add to each interval you are excluding.
Use this to add padding to the intervals specified using -XL. For example, '-XL 1:100' with a padding value of 20 would turn into '-XL 1:80-120'. This is typically used to add padding around targets when analyzing exomes.

int  0  [ [ -∞  ∞ ] ]


--interval-merging-rule / -imr

Interval merging rule for abutting intervals
By default, the program merges abutting intervals (i.e. intervals that are directly side-by-side but do not actually overlap) into a single continuous interval. However you can change this behavior if you want them to be treated as separate intervals instead.

The --interval-merging-rule argument is an enumerated type (IntervalMergingRule), which can have one of the following values:

ALL
OVERLAPPING_ONLY

IntervalMergingRule  ALL


--interval-padding / -ip

Amount of padding (in bp) to add to each interval you are including.
Use this to add padding to the intervals specified using -L. For example, '-L 1:100' with a padding value of 20 would turn into '-L 1:80-120'. This is typically used to add padding around targets when analyzing exomes.

int  0  [ [ -∞  ∞ ] ]


--interval-set-rule / -isr

Set merging approach to use for combining interval inputs
By default, the program will take the UNION of all intervals specified using -L and/or -XL. However, you can change this setting for -L, for example if you want to take the INTERSECTION of the sets instead. E.g. to perform the analysis only on chromosome 1 exomes, you could specify -L exomes.intervals -L 1 --interval-set-rule INTERSECTION. However, it is not possible to modify the merging approach for intervals passed using -XL (they will always be merged using UNION). Note that if you specify both -L and -XL, the -XL interval set will be subtracted from the -L interval set.

The --interval-set-rule argument is an enumerated type (IntervalSetRule), which can have one of the following values:

UNION
Take the union of all intervals
INTERSECTION
Take the intersection of intervals (the subset that overlaps all intervals specified)

IntervalSetRule  UNION


--intervals / -L

One or more genomic intervals over which to operate

List[String]  []


--inverted-read-filter / -XRF

Inverted (with flipped acceptance/failure conditions) read filters applied before analysis (after regular read filters).

List[String]  []


--lenient / -LE

Lenient processing of VCF files

boolean  false


--max-depth / -max-depth

Maximum read depth before a locus is considered poorly mapped
If the QC+ depth exceeds this value the site is considered to have EXCESSIVE_DEPTH

Integer  null


--max-depth-per-sample / -max-depth-per-sample

Maximum number of reads to retain per sample per locus. Reads above this threshold will be downsampled. Set to 0 to disable.

int  0  [ [ -∞  ∞ ] ]


--max-fraction-of-reads-with-low-mapq / -frlmq

If the fraction of reads at a base with low mapping quality exceeds this value, the site may be poorly mapped
If the number of reads at this site is greater than minDepthForLowMAPQ and the fraction of reads with low mapping quality exceeds this fraction then the site has POOR_MAPPING_QUALITY.

double  0.1  [ [ -∞  ∞ ] ]


--max-low-mapq / -mlmq

Maximum value for MAPQ to be considered a problematic mapped read
The gap between this value and mmq are reads that are not sufficiently well mapped for calling but aren't indicative of mapping problems. For example, if maxLowMAPQ = 1 and mmq = 20, then reads with MAPQ == 0 are poorly mapped, MAPQ >= 20 are considered as contributing to calling, where reads with MAPQ >= 1 and 20 are not bad in and of themselves but aren't sufficiently good to contribute to calling. In effect this reads are invisible, driving the base to the NO_ or LOW_COVERAGE states

int  1  [ [ 0  255 ] ]


--max-variants-per-shard

If non-zero, partitions VCF output into shards, each containing up to the given number of records.

int  0  [ [ 0  ∞ ] ]


--min-base-quality / -mbq

Minimum quality of bases to count towards depth
Bases with less than minBaseQuality are viewed as not sufficiently high quality to contribute to the PASS state

int  20  [ [ 0  255 ] ]


--min-depth / -min-depth

Minimum QC+ read depth before a locus is considered callable
If the number of QC+ bases (on reads with MAPQ > minMappingQuality and with base quality > minBaseQuality) exceeds this value and is less than maxDepth the site is considered PASS.

int  4  [ [ 0  ∞ ] ]


--min-depth-for-low-mapq / -mdflmq

Minimum read depth before a locus is considered a potential candidate for poorly mapped
We don't want to consider a site as POOR_MAPPING_QUALITY just because it has two reads, and one is MAPQ. We won't assign a site to the POOR_MAPPING_QUALITY state unless there are at least minDepthForLowMAPQ reads covering the site.

int  10  [ [ -∞  ∞ ] ]


--min-mapping-quality / -mmq

Minimum mapping quality of reads to count towards depth
Reads with MAPQ > minMappingQuality are treated as usable for variation detection, contributing to the PASS state.

int  10  [ [ 0  255 ] ]


--output / -O

Output file (BED or per-base format)

R GATKPath  null


--QUIET

Whether to suppress job-summary info on System.err.

Boolean  false


--read-filter / -RF

Read filters to be applied before analysis

List[String]  []


--read-index / -read-index

Indices to use for the read inputs. If specified, an index must be provided for every read input and in the same order as the read inputs. If this argument is not specified, the path to the index for each input will be inferred automatically.

List[GATKPath]  []


--read-validation-stringency / -VS

Validation stringency for all SAM/BAM/CRAM/SRA files read by this program. The default stringency value SILENT can improve performance when processing a BAM file in which variable-length data (read, qualities, tags) do not otherwise need to be decoded.

The --read-validation-stringency argument is an enumerated type (ValidationStringency), which can have one of the following values:

STRICT
LENIENT
SILENT

ValidationStringency  SILENT


--reference / -R

Reference sequence file

R GATKPath  null


--seconds-between-progress-updates / -seconds-between-progress-updates

Output traversal statistics every time this many seconds elapse

double  10.0  [ [ -∞  ∞ ] ]


--sequence-dictionary / -sequence-dictionary

Use the given sequence dictionary as the master/canonical sequence dictionary. Must be a .dict file.

GATKPath  null


--showHidden / -showHidden

display hidden arguments

boolean  false


--sites-only-vcf-output

If true, don't emit genotype fields when writing vcf file output.

boolean  false


--summary

Name of file for output summary
Callable loci summary counts will be written to this file.

R GATKPath  null


--tmp-dir

Temp directory to use.

GATKPath  null


--use-jdk-deflater / -jdk-deflater

Whether to use the JdkDeflater (as opposed to IntelDeflater)

boolean  false


--use-jdk-inflater / -jdk-inflater

Whether to use the JdkInflater (as opposed to IntelInflater)

boolean  false


--verbosity / -verbosity

Control verbosity of logging.

The --verbosity argument is an enumerated type (LogLevel), which can have one of the following values:

ERROR
WARNING
INFO
DEBUG

LogLevel  INFO


--version

display the version number for this tool

boolean  false


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GATK version 4.6.2.0 built at Sun, 13 Apr 2025 13:21:43 -0400.